I x T = P

This seems like a cryprtic formula, but it is utterly simple: Ideas x Time available = Priorities needed. When studying literature or when in contact with others ideas can spring to mind, not to mention the creative flow that can emerge from under the shower :-) Sometimes these ideas need time to take shape in a form that allows research, sometimes it is immediately clear that an idea is worthwhile to implement. The latter situation occurred this week when I was emailing to Mónica Vera about her application for a visiting scholarship to the Field Museum. When she was looking to make her proposal more specific, it immediately came to mind that making a revision of Plekocheilus would be a good idea that could serve several purposes: a) her application for a scholarship, b) the on-going research for making a checklist of Colombian land snails, c) our planned study of the Guayana Shield malacofauna and the speciation processes in that area. So, one more project added to the "wish list".

Fortunately, another project is nearing completion. My study of Orthalicidae from Ecuador is entering the reviewing phase. I will solicit the critical review of some colleagues before I submit the paper to a journal.

Making progress, steadily but slow

The focus in the Ecuadorian project is now on the analysis of the distributions. I decided first to pay more attention to the analysis of ecoregions and endemism. The latter concept is clearly contextual and I was tossing around with it when I found an interesting site that gave me some inspiration: the Andes-Amazon project in Peru and Bolivia. This project deals with plants, mammals, birds and amphibians, but in their methodology they also used Maxent modeling. The limited data on land mollusks (i.e. Orthalicidae) makes it necessary to tailor-made a definition on range-restricted species. I decided to limit it to those species known from a single locality. Those that are endemic to Ecuador deserve special attention, and when I plotted their localities three geographical clusters appeared. But clearly more research is needed before any definite conclusions can be drawn.
With my limited time (usually only during the weekends), progress is necessarily slow. So much to do and so little time...
Afbeelding 8

With some help and inspiration

The Maxent package I previously wrote about has a - like most freeware software - a discussion group which in my case was helpful to solve some initial problems. One problem - the grid output file that should be suitable for input in DIVA-GIS - was a bug in the beta version that I ran and was fixed very swiftly by the Steven Phillips, the main author. After this I could use the software to do useful analyses on the Ecuadorian data. The result is a series of distribution maps on (sub)generic level. Although several studies (e.g. Elith et al., 2006; Pearson et al., 2007) have shown that Maxent is robust enough to handle datasets with only a few occurrences per species, I preferred to do the analyses on aggregated data to get a more comprehensive result. Here is one of the maps, obtained by processing the grid output file of Maxent in DIVA-GIS.

ECU Plekocheilus maxent
Ecological niche model for Plekocheilus. The blue dots are localities where the genus is present. Colours represent logistic values of likelihood for occurrence: from 0-0.2 (dark green) to 0.8-1.0 (red).

When I out of curiousity tried to feed Maxent with occurrence data (also Plekocheilus) from the Guayana Shield, the resulting picture of the distribution showed a remarkable disjunct pattern. An inspirational indication that my hypothesis about linkages between the Pantepui region in Venezuela and the Andes in Colombia is worth further investigation!
VEN_GS_Plek_GS
Ecological niche model for Plekocheilus from Venezuela, Pantepui, as depicted by Maxent.
This even becomes more clear when all the known localities of Plekocheilus from Venezuela are taken into account.
VEN_Plekocheilus
Ecological niche model for all Plekocheilus known from Venezuela, as depicted by Maxent.

See also
this blog for comments on Maxent.

References:
Elith, J. e. a. (2006). Novel methods improve prediction of species' distributions from occurrence data. Ecography, 26, 129-151.
Pearson, R. G., C.J. Raxworthy, M. Nakamura & A. Townsend Peterson. (2007). Predicting species distributions from small numbers of occurrence records: a test case using cryptic geckos from Madagascar
. J. Biogeogr, 34, 102-117.